Ph.D. University of Maryland. Marine community assembly in shifting wetlands.
PhD and MSc, University of Jyväskylä, Finland. Studying the evolution of complex life cycle parasites
Ph.D. Oregon State University. Studying the ecology of trematodes in estuaries.
Faculty, University of Wisconsin
Faculty, Stanford University
Managing Editor, Integrated Environmental Assessment and Management
Research Scientist, Stanford University
Research Scientist, Smithsonian Tropical Research Institute, Panama.